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102

Rhodora

[June

showy flowers. Though it can hardly be considered
variety, it seems advisable to give it a distinct name.
Rhododendron

f. Harbisonii,

minus

forma

a botanical

nova.

A type differt floribus majoribus, 3.3-4 cm. longis in racemis cir
citer 10-floris, sepalis inaequalibus majoribus ovatis longe ciliatis.
Georgia:
Banks Co., May 18, 1911, T. G. Harbison (Nos. 615,
616 in Herb. Arnold Arboretum).
This differs from the typical form which has the flowers 2.5-3 cm.
the

long

and

and

larger

11

cm.

in

In

at

acuminate

leaves

and

that

but
very

the

somewhat

flowers

large
Gray,

the

Arnold

its

are

leaves

Chapmanii

not

acute,

re volute

the

in

7-8-flowered,

generally
also

length.
R.

resembles
or

racemes

racemes;

denser

densely

attaining
it

flower-clusters
has

species

flowers

larger

larger,
smaller

obtuse
and

beneath

lepidote

margin.

Arboretum.

THE

FORMS

OF PELTANDRA
Sidney

F.

VIRGINICA.

Blake.

Plate 94.
In
pp.

1836

in

Rafinesque,
elaborated

85-89,

his

the

New

Flora

of North

treatment

previous

America,

of the

genus

1,

part
Peltandra

into a monograph in which eight species were described, founded
principally on differences in leaf outline,1 although the number of
seeds
and

and

some

taken

recently

supposed

were

spathe

also

up

as

characters

diagnostic
Of

employed.
varieties

by

from

Rafinesque's

Mr.

Ivar

petiole,

species
on

Tidestrom,2

of field study of the plants in Maryland and Virginia.
last

fall

led

to

material
Botanical

of one
a

study
in

the
Club

or two
of

obviously

Rafinesque's

Gray
and

forms

Herbarium,
the

Boston

in

monograph
the

from

distinct

herbaria

Society

P. alba Raf. ( = P. sagittifolia (Mx.)
»Except
cospatha Raf.
2 Rhodora
12:47-50, pi. 83 (March, 1910).

Raf.),

of

connection

of Natural

were

the

basis

My collection

any

of the

scape,

two

New
History,

these

has

with

the

England
and

type of the subgenus

a

Leu

small
to

Forms of Peltandra

Blake,—

1912]

in

amount

characters

leaf
The

in their

forms

ears,

another

with

broad

of two

narrower

late

acuminate

The

two

which

from

differing

it has

been

other

apparently

each

and

leaves

with

leaves

treated

possible
in

short

one

broad

more

acute
forms

as

of

lobes

or

leaves
with

with
lanceo

or none

ears

var.

group

basal

of two
unequal

Tidestrom

of those

suggestive

groups:

and

short

composed

very

of by

three

narrower,

having
a third

having

strongly

into

naturally

forms

ears,

plants

of variation

range
fall

latifolia,

three

in

herbarium,

forms,

only.

forms,

of Sagittaria

longer,

own

my

seven

distinguish

103

virginica

at

all.

(Raf.)

heterophylla

Tidestrom and var. angustifolia (Raf.) Tidestrom make up this last
and

most

aberrant

acuminate
at

eared
and

P.

tapering

base.

These

angustifolia,

doubtedly
lead

basal

lobes,

ment

of the ears.

leaves
as

the

second

and

differing

the

subacute";
to

sufficient

marks

that

his

the

has

another

and

as distinct

of leaf

speci

but
and

narrow
develop

pointed

basal

lobes

often

twice

same

as

hastata

the

hastate

of

narrow

generally
P.

cordate

as long
Raf.,

acuminate,

has

not

the

name.
Rafinesque
"
is a Leucospatha

hence,

un

formae.

transitional
distinct

in size

of this

seeds,

equally

heterophyla1

with

be

adoption

several

one

possessing

oblong,

probability

P.

apparently

a form

possibly

length

justify

plant

few

divergent

may

in

to

themselves

among

usually

equal

close

lanceolate,
and

shortly

be maintained

of formae,

of these,

of leaf,

breadth

a

very

Rafinesque's

very
perhaps

pair

One

elongated

"leaves

lobes
me

to the

and

with

may

or

plants,

heterophyla

or narrowly

lance-oblong
rounded

two

intergrading,

mens

or

although

Peltandra

From

with

group,

leaves

however

seemed

to
re
like

the next," which is P. alba Raf. (now called P. sagittifolia (Mx.) Raf.),
the

of the

type

to whose

plant
Arum

well

as

virginicum

subgenus

distinguished

normal

the

leaves
understood

description
by

Elliott,

just

and

Raf.,

Leucospatha

well

quoted

considered

by

a

applies.

Rafinesque

identical with his P. hastata, is not by Elliott's description to be dis
from

tinguished
The

third

of the

plants

group,

ordinary

form

two

forms

comprising

with

hastate

beside

the

basal
type

lobes.
of

the

species, is distinguished from the last by its broad leaves with distinct
but
very
often

short

and

broad
subacute

broad

leaves
basal

ears.

nearly
lobes,

The

most

equilaterally
seems

to be

of these,

extreme
triangular,
identical

1 This orthography was apparently always employed
ous cases where this name was applied to new "species"

and
with

a form

with

obtuse

Rafinesque's

with
or
P.

by Raflnesque in the numer
in the "New Flora."

104

Rhodora
with

latifolia
long
one,

represented

Of the

broad."

has

mons,

broad

leaves

is here

more

leaf

acute

The

absence

the

differences

has

led
In

seem

me

the

taken

with

case

are

to rank

their

of the

type

of the

foliar

I

synopsis

forms,

and

have

elliptical
collected

foot

outline,
by

Com
ears

rounded

broadly
form,

which,

following
a narrower

has

species,

range,

or even

of a difference

with

coupled

that

as formae

plants

one

leaf

ears.

geographical

purely

natural

other

deltoid

of any

these

following

to be

narrower

and

tip

The

the

more
Delaware

a rounded

as

. .often

sagittate,..
with

from

sinus.

of a distinct

in the

in habitat,1

the

forms
sheet

with

closing

Tidestrom,

triangular

two

a single

by

or quite

nearly

broad

"leaves

and

[June

the
are

intergrades

rather

arranged

than
the

fact

that

frequent,

varieties.
forms

after

what

affiliations.

* Leaves
cm. wide), with distinct basal lobes 2-13 cm. long.
broad ( (3)7-17.5
+- Leaves
with blade!
cm. long, 8-17.5 cm. wide; basal lobes broad
(10)14-22
less than width of blade, deltoid
to
(3-7.5 cm. wide), short, nearly always
or rounded.
deltoid-lanceolate,
** Leaves
18.5-28
cm. across the
triangular,
very broad, almost equilaterally
or subacute,
sinus
tips of the ears, these obtuse
§ as long as main blade;
open.
1.
P.

nesque.
July

virginica

Peltandra
Raf.

latifolia
15,

1. c. 87.

"

(L.)
South

examined:
Specimens
1890
(Sereno
Watson).

f. latifolia

Kunth
New

Jersey
Massachusetts:
Delaware:

1890 (A. Commons).

and

n. comb.
(Raf.)
Rafi
Delaware,"
North
Cambridge,

Wilmington,

June

2,

*-* ** Leaves
cm. between
or elliptical-oblong,
not equilateral
(11-20
elliptical
tips of the ears).
= Leaves
cm. long, 16.5-17.5
cm. wide);
broad and rounded
16-19.5
(blade
or not projecting
at sides
of leaf,
lobes broadly
basal
rounded,
scarcely
sinus nearly or quite closed.
ï as long as the blade;
2.

P.

=

virginica

Delaware:

Herb.):

f. rotundata,

n. forma.3

Type

sheet

(in

Gray

Wilmington, June 2, 1890 (A. Commons).

= Leaves
more oblong (blade
cm. long, 8-17 wide);
basal lobes
(10)14-22
narrowed
toward
the tip, J-j the length of blade, usually
obtuse,
usually
sinus open.
somewhat
diverging;
3.

P.

virginica

Raf.,

1.

c.

87.

(typical
Typical

form).
specimens

P.

undvlata

examined:

Raf.,
Maine:

P.

canadensis
Livermore

1 Mr. Tidestrom found var. angustifolia growing with the type at Nanjemoy
Creek,
Dr. B. L. Robinson collected two forms under his number 161 at East
Maryland;
and I found three forms growing in a Sphagnum
bog in
Jaffrey, New Hampshire;
Canton, two about a pond in West Stoughton, and two in a ditch in woods in Norwood,
Massachusetts.
2 Blade is measured from top of petiole to tip of leaf.
3 P. viBGiNicA f. rotundata
n. forma, foliis rotundatis, lamina (vide supra) 16-19.5
lobis basilariis late rotundatis, vix aut omnino non
cm. longa, 16.5-17.5 cm. lata;
sinu prope vel ex toto clauso.
divergentibus, dimidio longis lamina;

Forms

Blake,—

1912]

of Peltandra

105

virginica

Falls, 1878 (Miss Furbish) ; Brunswick, 1882 (Miss Furbish) ; Spruce
E.
Pond, Cornish, July, 1900 (A. H. Norton). New Hampshire:

June
Reservation,
West
Stoughton,

Robinson
161 part).
Massachusetts:
II. Manning)-,
(IF.
26, 1894
Stony
May
17, 1895
(G. L. Chandler)-,
damp
ground
23 Sept.,
1911
(Blake
3378);
Sphagnum

and

(Ivar

Brook
near

L.

8 July,
1897
(B.
Brook
Reservation,

Jaffrey,
Beaver

pond,

bog, Canton, 13 Sept., 1911 (Blake 3325); ditch in woods, Norwood,
Delaware:
12 Oct., 1911 (Blake 3440).
Wilmington, June 2, 1890
Marshall Hall, July 23, 1890 (J. N.
(A. Commons). Maryland:
Rose)

June

1905

23,

Tidestrom)-,

swamp

Potomac

along

River, near Chainbridge, above Washington, July, 1904 (Ivar Tide
District

strom).

or

Columbia:

river

Washington,

May

swamp,

Miami River, Feb. 5, 1892 (J. II.
29, 1896 (E. S. Steele). Florida:
Simpson) ; swamps in water, vicinity of Eustis, Lake Co., April 15
30, 1894 (G. V. Nash 453).
•*— +- Leaves
cm. broad,
acute
to acuminate;
6-12
narrow,
comparatively
of blade),
narrow
basal
lobes longer (mostly one to two times the breadth
in
or linear lanceolate
(1-3.5 cm. broad near the middle),
mostly lanceolate
outline.
++ Leaves
cm. long, 6-12 cm. broad;)
larger and broader (main blade 10.5-23.5
cm. long, often twice the breadth of leaf, widely
basal lobes longer, (5)7-13
divaricate.
P.

4.

virginica

No.

sheet

4530,

Massachusetts,
same

in

data,

Cornish, July, 1890

Pond,

Spruce
setts:

n. forma.1
fP. hastata
Raf.
Type
f. hastifolia,
near pond,
West
my herb.;
ground
Stoughton,
damp
No.
Blake
23 Sept.,
paratype,
4531,
1911,
3377;
Maine:
Herb.
Other
examined:
specimens
Gray

Maiden,

2,

July

1890

(L.

L.

Connecticut:

Southington,

(no

other

++ Leaves
smaller
wide near middle);
divergent.
P.

1899

12,

L.

3441).

Andrews

(L.

data).

Hampshire:

1898 (W. Deane and B. L. Robinson 574).
abundant

Hampshire:

in shallow

8 July, 1897 (B. L. Robinson 161 part).
June

(II.

1911 (Blake

and narrower
10.5-20
(main blade
basal lobes shorter (2-7 cm. long),
n. forma.2
f. brachyota,
Contoocook
River,

virginica
New

Herb.):
New

July

1876

2,

July

ponds, Southington, July 12 and Aug. 18, 1899 (Bissell 881).

Michigan:

5.

in pond,

rare,

Massachu

Norton).

and

12 Oct.,

Moody)-, ditch in woods, Norwood,
820);

H.

(A.

Dame),

17,

1879

(H.

A.

Young)-

wet

cm. long, 3-5.5 cm.
the lobes only rarely

Type
East

sheet
Jaffrey,

(in Gray
6 Aug.,

Other specimens examined:

water

of river;

Massachusetts:

meadow

around

East

Jaffrey,

Lynnfield,
Doleful

Pond,

Stoneham, July 9, 1896 (W. P. Rich)-, sphagnum bog, Canton, 13

1 P. viRGiNiCA f. hastifolia, n. forma, foliis acutis vel acuminatis, lamina 10.5-23.5
cm. longis, latitudine
cm. longa, 6-12 cm. lata, lobis basilariis angustis, (5)7-13
laminae saepe duplo longioribus, late divaricatis.
2 P. viRQiNicA f. brachyota,
lamina
n. forma, foliis minoribus, angustioribus,
10.5-20 cm. longa, 3-5.5 cm. lata, auribus brevioribus, 2-7 cm. longis, raro divaricatis.

Rhodora

106

[June

Sept., 1911 (Blake 3328); shallow water, Centerville, July 3, 1900
(Mrs. C. I. Cheney); Stockbridge, Aug. 20, 1902 (R. Hoffman).
Delaware:
Wilmington, June 1890 (A. Commons).
* * Leaves

narrow
cm. wide),
with rounded,
(2.5-5
generally
acuminate,
or cordate
oblique
base, the ears when present
very short (1-2 cm. long)
and unequal.
+- Leaves
acute or acuminate,
15.5-19
cm. long, 4.5-5
oblong or lance-oblong,
cm. broad;
base oblique,
or unequally
and shortly eared.
rounded,
P.

6.
cording
"even

f. heterophylla

viRGiNiCA

Raf. 1. c. 88.
to

in

Philadelphia

Massachusetts:

sphagnum

3329

small,
[leaves
Maryland:
form]).

New

southern

Rafinesque,

near

n. comb.1

(Raf.)

P.

heterophyla

P. virginica var. heterophylla Tidesti >m, 1. c. 49.

only

the

Canton,

long

Marshall

Hall,

rare,

also
Delaware,
examined:
Specimens

Schuykill."

bog,
cm.
6.5-9

Ac

and

Jersey

13

1911

Sept.

perhaps
June

not
22,

really
1905

(Blake
this
(Ivar

Tidestrom).
*-

■*- Leaves
lanceolate,
long
very narrowly
cm. wide;
base oblique
or rounded.
2.5-3.5

acuminate,

23.5-28

cm.

long,

7. P. virginica f. angustifolia
(Raf.) n. comb. P. angustifolia
"
Vir
Raf. 1. c. 89. P. virginica var. angustifolia Tidestrom, 1. c. 50.
ginia

&c.,"

Nanjemoy

Rafinesque.

examined:

Specimens

Creek, June 1906 (Ivar Tidestrom).

Stoughton,

Maryland:

rare,

Massachusetts.

Explanation

of

Plate

94.

f. latifolia
Peltandra
Commons.
Not an ex
(Delaware,
virginica
Fig. 1.
treme of the form).
P. virginica f. rotundata (leaf from type sheet).
Fig. 2.
P. virginica f. hastifolia
(Michigan).
Fig. 3.
P. virginica f. brachyota (leaf from type sheet).
Fig. 4.
for March, 1910.)
(The three other forms were well figured by Mrs. Chase in Rhodoha
i It may be well to mention that the youngest leaves of the seedling (as shown by
sheets collected by Walter Deane on shore of "Glacialis,"
Cambridge, Massachusetts,
May 20, 1894) are of this type, suggesting that it represents the primitive leaf-form
in the species.

Plate

Rhodora

S. F. Blake del.
Formae

of

Peltandra

virginica.

94


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