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Cell Biology (Pollard) 2e .pdf



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S E C O N D

E D I T I O N

THOMAS D. POLLARD, MD
Sterling Professor, Department of Molecular, Cellular, and Developmental Biology
Yale University
New Haven, Connecticut

WILLIAM C. EARNSHAW, PhD, FRSE
Professor and Wellcome Trust Principal Research Fellow
Wellcome Trust Centre for Cell Biology, ICB
University of Edinburgh
Scotland, United Kingdom

WITH JENNIFER LIPPINCOTT-SCHWARTZ, PhD
Head, Section on Organelle Biology
Cell Biology and Metabolism Branch
National Institute of Child Health and Human Development
National Institutes of Health
Bethesda, Maryland

Illustrated by Graham T. Johnson

1600 John F. Kennedy Blvd.
Suite 1800
Philadelphia, PA 19103-2899

CELL BIOLOGY
SECOND EDITION
INTERNATIONAL EDITION

ISBN-13: 978-1-4160-2255-8
ISBN-10: 1-4160-2255-4
ISBN-13: 978-0-8089-2352-7
ISBN-10: 0-8089-2352-8

Copyright © 2008, 2004 by Thomas D. Pollard, William C. Earnshaw, Jennifer Lippincott-Schwartz:
Published by Elsevier Inc.
All rights reserved. No part of this publication may be reproduced or transmitted in any form or by
any means, electronic or mechanical, including photocopying, recording, or any information storage
and retrieval system, without permission in writing from the publisher. Permissions may be sought
directly from Elsevier’s Health Sciences Rights Department in Philadelphia, PA, USA: phone:
(+1) 215 239 3804, fax: (+1) 215 239 3805, e-mail: healthpermissions@elsevier.com. You may also
complete your request on-line via the Elsevier homepage (http://www.elsevier.com), by selecting
“Customer Support” and then “Obtaining Permissions.”

Notice
Knowledge and best practice in this field are constantly changing. As new research and
experience broaden our knowledge, changes in practice, treatment, and drug therapy may
become necessary or appropriate. Readers are advised to check the most current information
provided (i) on procedures featured or (ii) by the manufacturer of each product to be
administered, to verify the recommended dose or formula, the method and duration of
administration, and contraindications. It is the responsibility of the practitioners, relying on their
own experience and knowledge of the patients, to make diagnoses, to determine dosages and the
best treatment for each individual patient, and to take all appropriate safety precautions. To the
fullest extent of the law, neither the Publisher nor the Authors assume any liability for any injury
and/or damage to persons or property arising out of or related to any use of the material
contained in this book.
The Publisher
Library of Congress Cataloging-in-Publication Data
Pollard, Thomas D. (Thomas Dean), 1942–
Cell biology / Thomas D. Pollard, William C. Earnshaw; with Jennifer Lippincott-Schwartz ;
illustrated by Graham T. Johnson.—2nd ed.
p. cm.
Includes bibliographical references (p. ).
ISBN 1-4160-2255-4
1. Cytology. I. Earnshaw, William C. II. Title.
QH581.2.P65 2008
571.6—dc22
2006048515

Publishing Director: William Schmitt
Managing Editor: Rebecca Gruliow
Senior Developmental Editor: Jacquie Mahon
Publishing Services Manager: Joan Sinclair
Senior Book Designer: Ellen Zanolle
Marketing Manager: John Gore

Working together to grow
libraries in developing countries

Printed in China
Last digit is the print number:

www.elsevier.com | www.bookaid.org | www.sabre.org

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To Patty and Margarete and our families
The authors also express gratitude to their mentors, who helped to shape their views
of how science should be conducted. Tom Pollard thanks Sus Ito and Ed Korn for the
opportunity to learn microscopy and biochemistry under their guidance. He also
thanks Hugh Huxley and Ed Taylor for their contributions as role models, his former
colleagues at Johns Hopkins University for their insights regarding biophysics, and
Susan Forsburg for her help in the area of yeast biology. Bill Earnshaw thanks, in particular, Jonathan King, Stephen Harrison, Aaron Klug, Tony Crowther, Ron Laskey, and
Uli Laemmli, who provided a diverse range of incredibly rich environments in which
to learn that science at the highest level is an adventure that lasts a lifetime.

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Contributors
Jeffrey L. Corden, PhD

David Tollervey, PhD

Professor
Department of Molecular Biology and Genetics
Johns Hopkins Medical School
Baltimore, Maryland

Professor
Wellcome Trust Centre for Cell Biology
University of Edinburgh
Scotland, United Kingdom

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